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The consequences of these two main characteristics have been investigated in detail on an intertidal flat in the Schelde estuary in the Netherlands. The existence of a species minimum in the oligohaline section of the estuary, as postulated by Remane, could not be confirmed for nematodes and macrofauna for the Schelde, nor for several other European estuaries. Rather, a linear decrease in species richness from the sea to the freshwater was observed. The productivity of estuarine tidal flats depends on an in situ source of primary produced material, the microphytobenthos, as well as on the capacity of the macrofauna to transfer material from the water column to the sediments. 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