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The activity patterns of predators are highly variable over space and time and consequently prey are faced with the continual need to balance their habitat use (Chapter 7), foraging decisions (Chapter 2) and reproduction (Chapter 5) with the risk of predation (Lima & Dill 1990; Kats & Dill 1998; Lima 1998). Learning is the mechanism by which prey can achieve this important outcome because it allows prey to fine-tune their antipredator responses to variations in predation risk that can occur seasonally, across lunar cycles and on a moment-to-moment basis (Lima & Bednekoff 1999). 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