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I believe that there is robust evidence from host-specificity and from biogeography that allows us to look at the relative importance of co-evolutionary versus colonization events in determining host-parasite relationships. I am involved in the preparation of an interactive web-based key for the identification of sea lice (family Caligidae). These copepods are one of the major health problems for farmed fish worldwide. I am also interested in the colonization of marine and anchialine caves by crustaceans. These habitats seem to serve as refuges for basal representatives of numerous lineages and I have been studying the basal radiation of the Calanoida prior to their conquest of the open ocean pelagic zone.\r\n<p>\r\nI have been employed as a researcher at the Natural History Museum, London since 1974. I am also the Secretary of the Zoological Society of London, and I currently serve as a Trustee on the Board of SAHFOS, the Sir Alister Hardy Foundation for Ocean Science.","AbstractOtherLang":null,"AbstractLangCode":null,"AbstractLangID":null,"AutID":8785,"ND":"2003-10-29","UD":"2019-04-09","ORCID":"0000-0001-8170-7734"},"loaninfo":null,"pictures":[{"pic_id":119449,"Typ":3,"Descr":"Person on image","imis_show":1,"preferred":0,"URL":"/imis?page=image&pic=119449","ext":"jpg","url":"https://images.vliz.be/thumbs/119449_worms-steering-committee-meeting---innovocean-site-oostende-belgium-2016-06-16-front-row-from-left-to-right-tammy-horton-noc-nicole-boury-esnault-universit-d-aix-marseille-leen-vande.jpg"},{"pic_id":119459,"Typ":3,"Descr":"Person on image","imis_show":1,"preferred":0,"URL":"/imis?page=image&pic=119459","ext":"png","url":"https://images.vliz.be/thumbs/119459_worms-lifewatch-world-register-of-marine-cave-species-editor-workshop-at-innovocean-site-oostende-belgium-2016-02-22-25-from-left-to-right-alejandro-mart-nez-garc-a-meg-ise-fernando-lvarez-n.png"},{"pic_id":124843,"Typ":3,"Descr":"Person on image","imis_show":1,"preferred":0,"URL":"/imis?page=image&pic=124843","ext":"jpg","url":"https://images.vliz.be/thumbs/124843_from-left-to-right-shokoofeh-shamsi-geoff-boxshall-rod-bray-robert-adlard-david-gibson-haylee-weaver-stefanie-dekeyzer-janine-caira-sarah-perche-malin-strand-kerry-hadfield-alfonso-herrer.jpg"}],"institutes":[{"instituterec":{"OrderNr":1,"Acronym":null,"ENFunction":"Merit Researcher, Invertebrates Division","InsIDtmp":1941,"OrigNameLangCode":null,"OrigNameLangID":null,"FullOrigName":null,"InsID":1941,"Function":"Merit Researcher, Invertebrates Division","BeginDay":null,"BeginMonth":null,"BeginYear":null,"Begindate":"","Enddate":"","PIAdrID":121087,"AdrID":113072,"Line1":null,"Line2":null,"Line3":null,"Line4":null,"Phone":"+44-(0)20-7942 57 49","GSM":null,"Email":"g.boxshall@nhm.ac.uk","EnvName":"UK","EncAddress":"Cromwell Road, London, SW7 5BD, UK","FullStandardName":"Natural History Museum; 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Bouchet, P.; Boxshall, G.; Arvanitidis, C.; Appeltans, W. (2026). European Register of Marine Species. Accessed at http://www.marbef.org/data/erms.php on yyyy-mm-dd","Title":"European Register of Marine Species version 2","DasType":"Data"},{"DasID":5399,"Acronym":null,"Citation":"Türkay, M. (2015). Personal Decapoda distribution Database for Europe.","Title":"Personal Decapoda distribution Database for Europe","DasType":"Data"},{"DasID":5276,"Acronym":"WoRCS","Citation":"Gerovasileiou, V.; Martínez García, A.; Álvarez Noguera, F.; Boxshall, G.; Humphreys, W.F.; Jaume, D.; Becking, L.E.; Muricy, G.; van Hengstum, P.J.; Yamasaki, H.; Bailly, N.; Iliffe, T.M. (2026). World Register of Marine Cave Species (WoRCS). Accessed at https://www.marinespecies.org/worcs on yyyy-mm-dd","Title":"World Register of marine Cave Species","DasType":"Data"},{"DasID":4165,"Acronym":null,"Citation":"Boxshall, G.A. (2026). World List of Bochusacea. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&amp;id=391550 on yyyy-mm-dd","Title":"World List of Bochusacea","DasType":"Data"},{"DasID":4166,"Acronym":null,"Citation":"Boxshall, G.A. (2026). World List of Brachypoda. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&amp;id=238273 on yyyy-mm-dd","Title":"World List of Brachypoda","DasType":"Data"},{"DasID":4184,"Acronym":null,"Citation":"Boxshall, G. (2026). World List of Merostomata. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&amp;id=150507 on yyyy-mm-dd.","Title":"World List of Merostomata","DasType":"Data"},{"DasID":4187,"Acronym":null,"Citation":"Boxshall, G.A. (2026). World List of Mystacocarida. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&amp;id=1079 on yyyy-mm-dd","Title":"World List of Mystacocarida","DasType":"Data"},{"DasID":4207,"Acronym":null,"Citation":"Boxshall, G.A.; Huys, R.; Walter, T.C. (2026). World List of Tantulocarida. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&amp;id=1083 on yyyy-mm-dd","Title":"World List of Tantulocarida","DasType":"Data"},{"DasID":3104,"Acronym":null,"Citation":"Walter, T.C.; Boxshall, G. (2026). World of Copepods Database. Accessed at https://www.marinespecies.org/copepoda on yyyy-mm-dd","Title":"World of Copepods","DasType":"Data"}],"references":{"A1":[{"BRefID":408363,"RR":"<b>Bernot, J.P.; Khodami, S.; Boyen, J.; De Troch, M.; Boxshall, G.A.; Arbizu, P.M.</b> (2025). Copepod phylogenomics supports Canuelloida as a valid order separate from Harpacticoida. <i>Mol. Phylogenet. Evol. 206</i>: 108311. <a href=\"https://dx.doi.org/10.1016/j.ympev.2025.108311\" target=\"_blank\">https://dx.doi.org/10.1016/j.ympev.2025.108311</a>","AutID":10636,"MonDate":null,"AnaDate":2025,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":368380,"RR":"<b>Boxshall, G.A.</b> (2023). Resolving taxonomic and nomenclatural problems in the genus <i>Caligus</i> O.F. Müller, 1785 (Copepoda: Caligidae). <i>Zootaxa 5360(4)</i>: 545-567. <a href=\"https://dx.doi.org/10.11646/zootaxa.5360.4.5\" target=\"_blank\">https://dx.doi.org/10.11646/zootaxa.5360.4.5</a>","AutID":10636,"MonDate":null,"AnaDate":2023,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":325398,"RR":"<b>Kim, I.-H.; Boxshall, G.A.</b> (2020). A revision of the family Archinotodelphyidae Lang, 1949 (Copepoda: Cyclopoida: Oithonida), with the recognition of 15 new species</strong>. <i>Zootaxa 4801(1)</i>: 1-56. <a href=\"https://dx.doi.org/10.11646/zootaxa.4801.1.1\" target=\"_blank\">https://dx.doi.org/10.11646/zootaxa.4801.1.1</a>","AutID":306454,"MonDate":null,"AnaDate":2020,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":311146,"RR":"<b>Bernot, J.P.; Boxshall, G.A.</b> (2019). Two new species of parasitic copepods from the genera <i>Nothobomolochus</i> and <i>Unicolax</i> (Cyclopoida: Bomolochidae) from Australian waters. <i>PeerJ 7</i>: e6858. <a href=\"https://dx.doi.org/10.7717/peerj.6858\" target=\"_blank\">https://dx.doi.org/10.7717/peerj.6858</a>","AutID":10636,"MonDate":null,"AnaDate":2019,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":294248,"RR":"<b>Boxshall, G.</b> (2018). The sea lice (Copepoda: Caligidae) of Moreton Bay (Queensland, Australia), with descriptions of thirteen new species. <i>Zootaxa 4398(1)</i>: 1-172. <a href=\"https://dx.doi.org/10.11646/zootaxa.4398.1.1\" target=\"_blank\">https://dx.doi.org/10.11646/zootaxa.4398.1.1</a>","AutID":8785,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":293145,"RR":"<b>Boyko, C.B.; Boxshall, G.</b> (2018). Recognition of Cumoniscidae Nierstrasz & Brender à Brandis, 1923, as the senior subjective synonym of Deoterthridae Boxshall & Lincoln, 1987 (Crustacea: Tantulocarida). <i>Proc. Biol. Soc. Wash. 131(1)</i>: 61-64. <a href=\"https://dx.doi.org/10.2988/17-00029\" target=\"_blank\">https://dx.doi.org/10.2988/17-00029</a>","AutID":8785,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":304036,"RR":"<b>Ohtsuka, S.; Nishikawa, J.; Boxshall, G.A.</b> (2018). A new species of <i>Peniculus</i> (Copepoda: Siphonostomatoida) parasitizing mesopelagic myctophid fish: first discovery of colonization of the genus in deep water. <i>Parasite 25</i>: 58. <a href=\"https://dx.doi.org/10.1051/parasite/2018057\" target=\"_blank\">https://dx.doi.org/10.1051/parasite/2018057</a>","AutID":10636,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":290080,"RR":"<b>Özak, A.A.; Rodrigues, A.M.V.; Vieira, N.S.S.; Rosa, R.M.G.; Yanar, A.; Koyuncu, E.; Boxshall, G.A.</b> (2018). <i>Lepeophtheirus azoricus</i> n. sp. (Copepoda: Caligidae) parasitic on dusky grouper, <i>Epinephelus marginatus</i>, from Atlantic waters off the Azores, Portugal. <i>Mar. Biodiv. 48(2)</i>: 1045-1055. <a href=\"https://dx.doi.org/10.1007/s12526-017-0797-x\" target=\"_blank\">https://dx.doi.org/10.1007/s12526-017-0797-x</a>","AutID":10636,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":296596,"RR":"<b>Özak, A.A.; Ternengo, S.; Boxshall, G.A.</b> (2018). Redescription of <i>Caligus pagelli</i> Delamare Deboutteville & Nuñes-Ruivo, 1958 (Copepoda: Caligidae) on the common pandora, <i>Pagellus erythrinus</i> (Linnaeus), in western Mediterranean waters off Corsica, France. <i>Syst. Parasitol. 95(6)</i>: 591-601. <a href=\"https://dx.doi.org/10.1007/s11230-018-9799-8\" target=\"_blank\">https://dx.doi.org/10.1007/s11230-018-9799-8</a>","AutID":10636,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":300787,"RR":"<b>Özak, A.A.; Yanar, A.; Sakarya, Y.; Boxshall, G.A.</b> (2018). The discovery of <i>Lepeophtheirus acutus</i> Heegaard, 1943 (Copepoda: Caligidae) from two new elasmobranch hosts in the Mediterranean Sea, and a comparative redescription of <i>Lepeophtheirus rhinobati</i> Luque, Chaves et Cezar, 1998. <i>Acta Parasitol. 63(3)</i>: 454-473. <a href=\"https://dx.doi.org/10.1515/ap-2018-0055\" target=\"_blank\">https://dx.doi.org/10.1515/ap-2018-0055</a>","AutID":10636,"MonDate":null,"AnaDate":2018,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":221925,"RR":"<b>Costello, M.J.; Bouchet, P.; Boxshall, G.; Fauchald, K.; Gordon, D.; Hoeksema, B.W.; Poore, G.C.B.; Van Soest, R.W.M.; Stöhr, S.; Walter, T.C.; Vanhoorne, B.; Decock, W.; Appeltans, W.</b> (2013). Global coordination and standardisation in marine biodiversity through the World Register of Marine Species (WoRMS) and related databases. <i>PLoS One 8(1)</i>: 20 pp. <a href=\"https://dx.doi.org/10.1371/journal.pone.0051629\" target=\"_blank\">https://dx.doi.org/10.1371/journal.pone.0051629</a>","AutID":8785,"MonDate":null,"AnaDate":2013,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":220199,"RR":"<b>Appeltans, W.; Ahyong, S.T.; Anderson, G.; Angel, M.V.; Artois, T.; Bailly, N.; Bamber, R.N.; Barber, A.; Bartsch, I.; Berta, A.; Blazewicz-Paszkowycz , M.; Bock, P.; Boxshall, G.; Boyko, C.B.; Nunes Brandão, S.; Bray, R.A.; Bruce, N.L.; Cairns, S.D.; Chan, T.-Y.; Cheng, L.; Collins, A.G.; Cribb, T.H.; Curini-Galletti, M.; Dahdouh-Guebas, F.; Davie, P.J.F.; Dawson, M.N.; De Clerck, O.; Decock, W.; De Grave, S.; de Voogd, N.J.; Domning, D.P.; Emig, C.C.; Erséus, Ch.; Eschmeyer, W.; Fauchald, K.; Fautin, D.G.; Feist, S.W.; Fransen, C.H.J.M.; Furuya, H.; García-Alvarez, O.; Gerken, S.; Gibson, D.; Gittenberger, A.; Gofas, S.; Gómez-Daglio, L.; Gordon, D.P.; Guiry, M.D.; Hernandez, F.; Hoeksema, B.W.; Hopcroft, R.R.; Jaume, D.; Kirk, P.; Koedam, N.; Koenemann, S.; Kolb, J.B.; Kristensen, R.M.; Kroh, A.; Lambert, G.; Lazarus, D.B.; Lemaitre, R.; Longshaw, M.; Lowry, J.; MacPherson, E.; Madin, L.P.; Mah, C.; Mapstone, G.; McLaughlin, P.A.; Mees, J.; Meland, K.; Messing, C.G.; Mills, C.E.; Molodtsova, T.N.; Mooi, R.; Neuhaus, B.; Ng, P.K.L.; Nielsen, C.; Norenburg, J.; Opresko, D.M.; Osawa, M.; Paulay, G.; Perrin, W.; Pilger, J.F.; Poore, G.C.B.; Pugh, Ph.R.; Read, G.B.; Reimer, J.D.; Rius, M.; Rocha, R.M.; Sáiz Salinas, J.I.; Scarabino, V.; Schierwater, B.; Schmidt-Rhaesa, A.; Schnabel, K.E.; Schotte, M.; Schuchert, P.; Schwabe, E.; Segers, H.; Self-Sullivan, C.; Shenkar, N.; Siegel, V.; Sterrer, W.; Stöhr, S.; Swalla, B.; Tasker, M.L.; Thuesen, E.V.; Timm, T.; Todaro, M.A.; Turon, X.; Tyler, S.; Uetz, P.; van der Land, J.; Vanhoorne, B.; van Ofwegen, L.P.; Van Soest, R.W.M.; Vanaverbeke, J.; Walker-Smith, G.K.; Walter, T.C.; Warren, A.; Williams, G.C.; Wilson, S.P.; Costello, M.J.</b> (2012). The magnitude of global marine species diversity. <i>Curr. Biol. 22(23)</i>: 2189-2202. <a href=\"http://dx.doi.org/10.1016/j.cub.2012.09.036\" target=\"_blank\">http://dx.doi.org/10.1016/j.cub.2012.09.036</a>","AutID":8785,"MonDate":null,"AnaDate":2012,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":244143,"RR":"<b>El-Rashidy, H.H.; Boxshall, G.A.</b> (2012). A new copepod (Siphonostomatoida: Lernanthropidae) parasitic on a Red Sea immigrant dragonet (Actinopterygii: Callionymidae), with a review of records of parasitic copepods from dragonets. <i>Syst. Parasitol. 81(2)</i>: 87-96. <a href=\"http://dx.doi.org/10.1007/s11230-011-9326-7\" target=\"_blank\">http://dx.doi.org/10.1007/s11230-011-9326-7</a>","AutID":306454,"MonDate":null,"AnaDate":2012,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":288533,"RR":"<b>Hayes, P.; Justine, J.-L.; Boxshall, G.A.</b> (2012). The genus <i>Caligus</i> Müller, 1785 (Copepoda: Siphonostomatoida): two new species from reef associated fishes in New Caledonia, and some nomenclatural problems resolved. <i>Zootaxa 3534(3534)</i>: 21-39","AutID":10636,"MonDate":null,"AnaDate":2012,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":287945,"RR":"<b>Suárez-Morales, E.; Boxshall, G.A.</b> (2012). A new species of <i>Sabellacheres</i> M. Sars, 1862 (Copepoda: Gastrodelphyidae) from a deep-water benthic polychaete in Antarctic waters, with a key to the species of the genus. <i>Syst. Parasitol. 83(1)</i>: 65-75. <a href=\"https://dx.doi.org/10.1007/s11230-012-9371-x\" target=\"_blank\">https://dx.doi.org/10.1007/s11230-012-9371-x</a>","AutID":10636,"MonDate":null,"AnaDate":2012,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":137915,"RR":"<b>Brugnano, C.; Boxshall, G.A.; Costanzo, G.; Zagami, G.</b> (2009). Morphological variation and a remarkable new sexual dimorphism in the <i>Pseudocyclops xiphophorus</i> population of Lake Faro in northeastern Sicily. <i>Sci. Mar. (Barc.) 73(2)</i>: 347-355. <a href=\"https://dx.doi.org/10.3989/scimar.2009.73n2347\" target=\"_blank\">https://dx.doi.org/10.3989/scimar.2009.73n2347</a>","AutID":312604,"MonDate":null,"AnaDate":2009,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":254242,"RR":"<b>Costello, M.J.; Boxshall, G.A.; Boyko, C.B.; Høeg, J.T.; Markham, J.; Appeltans, W.</b> (2009). Development of sustainable authoritative online species databases. <i>Integrative and Comparative Biology 49</i>: E217-E217","AutID":10636,"MonDate":null,"AnaDate":2009,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":254243,"RR":"<b>Costello, M.J.; Boxshall, G.A.; Boyko, C.B.; Høeg, J.T.; Markham, J.; Appeltans, W.</b> (2009). How can we best make authoritative biological information available online? <i>Integrative and Comparative Biology 49</i>: E38-E38","AutID":10636,"MonDate":null,"AnaDate":2009,"PeerRev":1,"outputType":"1_A1","OpenAcc":1},{"BRefID":244119,"RR":"<b>El-Rashidy, H.H.; Boxshall, G.A.</b> (2009). Parasites gained: Alien parasites switching to native hosts. <i>J. Parasitol. 95(6)</i>: 1326-1329. <a href=\"http://dx.doi.org/10.1645/GE-2190.1\" target=\"_blank\">http://dx.doi.org/10.1645/GE-2190.1</a>","AutID":293595,"MonDate":null,"AnaDate":2009,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":244124,"RR":"<b>Di Capua, I.; Boxshall, G.A.</b> (2008). Records of Oncaeidae (Copepoda) from the Gulf of Naples, with new records of three species of <i>Triconia</i>. <i>J. Mar. Biol. Ass. U.K. 88(7)</i>: 1407-1415. <a href=\"http://dx.doi.org/10.1017/S002531540800194X\" target=\"_blank\">http://dx.doi.org/10.1017/S002531540800194X</a>","AutID":10636,"MonDate":null,"AnaDate":2008,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":105102,"RR":"<b>Jaume, D.; Boxshall, G.A.; Bamber, R.N.</b> (2006). A new genus from the continental slope off Brazil and the discovery of the first males in the Hirsutiidae (Crustacea: Peracarida: Bochusacea). <i>Zool. J. Linn. Soc. 148(2)</i>: 169-208. <a href=\"https://dx.doi.org/10.1111/j.1096-3642.2006.00235.x\" target=\"_blank\">https://dx.doi.org/10.1111/j.1096-3642.2006.00235.x</a>","AutID":10636,"MonDate":null,"AnaDate":2006,"PeerRev":1,"outputType":"1_A1","OpenAcc":0},{"BRefID":6458,"RR":"<b>Boxshall, G.A.; Ohtsuka, S.</b> (2001). Two new families of copepods (Copepoda: Siphonostomatoida) parasitic on echinoderms. <i>J. Crust. 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