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In this framework, the overall aim of BIOFUN is to characterise, through an ecosystemic approach, two deep-sea habitats – the mid-slope and abyssal plain – to understand the linkages between biodiversity patterns and ecosystem functioning in relation to environmental conditions along a trophic gradient, from Eastern Atlantic to the Western, Central and Eastern Mediterranean, enabling the simulation of their potential response to changing trophic conditions. This is the first proposal aiming at a complete investigation of the entire food web, from viruses and microbes to megafauna, including commercial species. In particular, BIOFUN is structured in 4 major work-packages: 1) physical and geochemical habitat conditions; 2) community structure: biodiversity and biogeography; 3) ecosystem functioning: food web processes and life-history patterns; and 4) linkages between ecosystem functioning and biodiversity: tools for disturbance evaluation. BIOFUN is a consortium of 10 European partners leading deep-sea research at the international level and with wide expertise in the use of large platforms and state-of-the art methods. 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